Unlocking each skill or skill tree is gained by meeting the requirements. Once you have all listed requirements done for the Ixi in question, you don't need to do anything more, the Ixi will already have the new skills available!
Some skills or skill trees are obtained by completing Story Modes, using items, and other more miscellaneous methods. You can find these under any tab other than Physical. Here's a quick rundown on what skills can be found in which tab, for your convenience in finding things.
Physical skills are obtained naturally by leveling up!
Radiation skills are obtained through Story Modes and in one case, via Chasmjumping.
Fridge Parasite - Skill: Fridge Werewolf: Can appear in Fridge Werewolf alternate form for EXP and rewards. Consuming 5 food items in one comment has a 25% chance of spawning an Inert Parasite item. (Can stack if more than 5 items are used in one commment, can only spawn up to 3 Inert Parasites/week per Fridge Werewolf)
Mymber's Heart Valve - Special Ability: Mymber Valve Transplant: Nullifies one chosen Radiation Sickness in a Rad Sick Ixi. If a Mymber is equipped, that Mymber cannot nullify a second Rad Sickness.
In both non-Rad Sick and Rad Sick Ixi, has a 30% chance of temporarily nullifying Radiation Sickness effects in other Ixi in certain situations: in Breeding, this Ixi has a 30% chance to deactivate Radiation Sicknesses that give breeding debuffs (Infertility, Recessive, Fever Dreamer, etc) in the other partner. In activities, if depicted with another Ixi that happens to have a Rad Sickness that can impact their roll, there is a 30% chance this Ixi can nullify their Rad Sick traveling companion's ailment for that roll.
This ability also grants a resistance to certain Temporary Conditions - this resistance applies to both the owner of the Mymber's Heart Valve as well as any other Ixi depicted in the piece.
Poison Blade - Special Ability: Brutal Rending: The Ixi holding this blade will always roll more than one item in Hunting and Combat [unless affected by radiation sickness]!
Witch's Brew Tea - Special Ability: Forgive and Forget: The Ixi with this ability may reroll one exploration, hunting, mining, or combat roll.
Items can grant single skills or entire skill trees. Certain items can be equipped to other items/companions.
Barghest - Companion Ability: Barghest: Adds a bonus +50 Attack and +50 Defense in Hunting. (OVERRIDES Radiation Sickness if present!)
When entering Legendary Hunting with a party of 3 or Less Rixixi, a Barghest can count as an additional Rixixi for the party - Barghests have a stat pool of 150/150/100/100 for Legendary Hunting. This bonus cannot come into play if there are 4 Ixi in the party already.
Bombardier - Companion Ability: Bombardier: Makes the boost from superhot peppers (Bhut Jolokia, Magmatic Spitfire, Incendiary Witchcraft, and Wall Crawler) permanent.
Legendary Companion Second Ability:
Bombardier: Fire-Breather: Increases elemental attacks' damage by 150%. Counts for null and chasm as well as regular radiation.
Battle Sword Special Ability: Honed Edge: Rolls any Attack stat rolls in combat twice, with the better roll being used.
Bloodletting Dagger Special Ability: Ritual: If sacrificing a Rixixi at the Phoenix Shrine, link this Rixixi to use their Bloodletting Dagger as the blade. Bloodletting Dagger has a 75% chance to kill a Rixixi. If it does kill the sacrifice, all stats are removed from the sacrifice and absorbed into the holder of the Bloodletting Dagger. If it does not kill the sacrifice, half of the sacrifice's stats are removed as usual and absorbed into the owner of the Bloodletting Dagger. Unuploaded genos still have a 100% death rate, but are considered to have base stats.
Caverndrake - Companion Ability: Caverndrake: Adds a bonus +100 Defense in Hunting and Mining.
If the Rixixi has a Tunneldrake equipped, the Tunneldrake's return is doubled.
Legendary Companion Second Ability:
Caverndrake: Death From Below: When Legendary Hunting, adds 15% Success chance to entire party. Adds Pet Treat effect to roll (35% chance of luring a random companion). The Rixixi with this companion cannot use Pet Treats during Legendary Hunting as a result.
Couler - Companion Ability: Couler: If the sun is depicted or described as being out in the art for the roll, This companion can add on an entire additional roll (including seasonal item returns) to its companion Rixixi whenever the Rixixi it is attached to is depicted in any loot-dropping activity! The additional roll will be rolled with base stats rather than the Rixixi's stats, as the Couler is picking up the items rather than the Rixixi.
Crow - Companion Ability: Crow: 15% chance of dodging any attack in combat through sheer intimidation. Passive, does not count as equipment.
Legendary Companion Second Ability:
Crow: Scarecrow: In any loot-dropping activity, the crow returns a full Hunting roll vs. Plant Creature prey at base stats.
Deathtrap - Companion Ability: Deathtrap: Doubles the Rixixi companion's contributions to raids.
Legendary Companion Second Ability:
Deathtrap: Suneater: Automatically and passively counters 10% of any elemental radiation damage used in combat.
Can be used once per combat to counter 150% of any elemental radiation damage used that turn. Suneater is used as the Rixixi's active skill in this case. If no elemental radiation skills are used this turn, Suneater is still counted as used for that combat encounter.
Drakescale Armor Special Ability: Heavy Armor: Automatically blocks damage in combat as long as equipped items are allowed - Damage blocked = [Defense/2]+[Special/4]+4d100. IF your Defense is above 200, 10% chance to automatically block one incoming attack per round (defaults to the strongest attack). Decreases Priority by 100 if Attack stat is under 300.
Fallen Leaftail Gecko - Companion Ability: Fallen Leaftail Gecko: The gecko glides overhead to seek a treasure out. 50% chance to roll any one specified common or uncommon item in exploring or mining. Item will be rolled at base stats. Specified item must be one of the already-available items for the roll. If no item is listed or the listed item is incompatable with the roll, this part of Leaftail Gecko's skill will not be rolled.
Also acts as a version of the Pumpkin Spice Apple Cider boon that can be activated on any loot-dropping roll the attached Rixixi is in: Fallen Leaftail Gecko doubles all seasonal returns! In Karma Events, +5 Positive or Negative Karma will be added to the Rixixi with this companion upon activation of this skill (please list whether you want positive or negative karma!) Pumpkin Spice Apple Cider cannot be used on this Rixixi as it would have the same effect.
Geode Mimic - Companion Ability: Geode Mimic: When equipped to a Rixixi, the user can feed 2 gemstones of any rarity to the Geode Mimic to receive 1 randomized gemstone in return! Use this ability in the Player Inventories journal under "Use An Item" to activate, and link the Rixixi with this companion. This ability can be activated up to 5 times per week.
Grand Old Rooter - Companion Ability: Grand Old Rooter: Adds a bonus +100 Defense in Hunting and a bonus +100 Speed in Exploring. (OVERRIDES Radiation Sickness if present!) In Combat, AUTOMATICALLY defends for [2d100] (Acts as armor.)
Legendary Companion Second Ability:
Grand Old Rooter: Keeping the Herd Together: If the Rixixi this Legendary Companion is attached to has a Grand Old Rooter (including this one), a Rooter, and a Mini Rooter, all Rooters of any type (including Rooters owned by other Rixixi in the image) count as Rixixi when lowering a Legendary Hunting failure chance, and they count towards the final stat pool (Grand Old Rooters have 250/350/200/100, Rooters have 100/150/100/50, and Mini Rooters have 50/50/50/50).
Graverobber - Companion Ability: Graverobber: 50% chance of dropping a Death-Charged Fogshard in any loot-dropping activity.
Legendary Companion Second Ability:
Graverobber: Soft Earth: Can be used during combat and will not take up the Rixixi's combat turn (Soft Earth can be used as an extra skill during combat).
Spends one turn digging underground, shielding their Rixixi for 20% of damage due to dirt being kicked up. The next turn, they attack for [2d20+(attack*2)]!
Horseman Heart Special Ability: Horseman Host: Any negative effects from Radiation Sickness are inverted. If they cannot be inverted, they simply will not trigger. Does not effect Chasmhome or any Coat radiation sicknesses.
Holo Hacker - Companion Ability: Holo Hacker: Whenever the Rixixi it is attached to is depicted in a loot-dropping activity, adds on an entire additional roll, rolled at base stats. Adds Streetwise to the additional roll even if the Rixixi doesn't have it - only to the additional roll if the Rixixi doesn't have it. Additionally, during Karma events only, for every contraband item found, 1 Negative Karma is added. The Holo Hacker knows what those are for.
Holo Helper - Companion Ability: Holo Helper: Whenever the Rixixi it is attached to is depicted in a loot-dropping activity, adds on an entire additional roll, rolled at base stats. Adds Streetwise to the additional roll even if the Rixixi doesn't have it - only to the additional roll if the Rixixi doesn't have it. Additionally, during Karma events only, for every contraband item found, 1 Positive Karma is added. The Holo Helper hopes you're cleaning it off the street.
Hunting Knife Special Ability: Easy Pelts: The Rixixi with this item equipped, while hunting, will also roll one extra item - rolled from pelt-type items such as leather, scales, shells, and so on.
Kaurphyt Raven - Companion Ability: Kaurphyt Raven: This companion adds a bonus +50 to Special to its companion Rixixi in Explorations and Combat (OVERRIDES Radiation Sickness if present!), and protects Rixixi companion by increasing success chance in Danger Chasmjumping Rolls by 15%.
Kellas Cat - Companion Ability: Kellas Cat: This companion adds a bonus +50 to Attack to its companion Rixixi in Hunting and Combat (OVERRIDES Radiation Sickness if present!), and may go out on adventures on its own - check in with your cat to see if it's brought anything interesting home. (Use this secondary ability in the Kellas Cat prompt to activate, and link the Rixixi with this companion. This ability can be activated once a week.)
Kyaine - Companion Ability: Kyaine: Nullifies one chosen radiation sickness if present. This stacks with Mymber.
However, any O'urtnx NPCs cannot assist this Rixixi.
Legendary Companion Second Ability:
Kyaine: Null Field: Radiation skills other than Null Radiation cannot be used by opponents or companion Rixixi.
Leather Armor Special Ability: Light Armor: Automatically blocks damage in combat as long as equipped items are allowed - Damage blocked = ([Defense/4]+[Speed/2])/2d6. IF your Speed is above 200, 10% chance to automatically dodge the next attack after your turn. Increases Priority by 50 if Speed stat is over 300.
Lemon Pig - Companion Ability: Lemon Pig: 80% chance of digging up an uncommon item in Explorations and Mining. Returns 1 Citrus Fruit on failed roll.
Lemon Pigs count as Mini Rooters for the Grand Old Rooter's ability.
Leyline Barghest - Companion Ability: Leyline Barghest: As they are closely related to the Barghest Companion, Leyline Barghests have the original Barghest skill and an additional Leyline-related skill.
Leyline Barghest (Extra Skill): When equipped to a living Rixixi, treats that Rixixi as a Ghost in all ways except for genetics.
Leyline Kaurphyt - Companion Ability: Leyline Kaurphyt: When equipped to a Rixixi, the Leyline Kaurphyt can cure one Temporary Condition once a week! Use this ability in the Player Inventories journal under "Use An Item" to activate, and link the Ixi with this companion! Leyline Kaurphyts are not limited to curing their companion Ixi, and can heal other users' Ixi with that player's permission! This counts as curing a temporary condition with items for Medic specialty.
Additionally, when equipped to a living Rixixi, treats that Rixixi as a Ghost in all ways except for genetics.
Leyline Mymber - Companion Ability: Leyline Mymber:
Counts as a Mymber in any scenarios that specifically require one, such as needing a tutor for the Mymber Tome. MymbMumbles Lookalikes are Mymbers with an extra skill.
Leyline Mymber: The Rixixi with this companion will obtain an extra item in loot-dropping activities if depicted with water (includes ice and snow)! Also, the Rixixi counts as a Ghost in any situations where this would alter dialogue - this does not add the +Ghost gene, the Rixixi is essentially able to see ghosts and is respected by ghosts!
Leyline Rooter - Companion Ability: Leyline Rooter: In Explorations and Mining, will dig up any of the following items: Slab of Raw Meat, Flesh Slab, Teeth and Claws, Leather, Death-Charged Fogshard*, Mymber Heart*, B-NOMALY Greymatter Sample*
*Contraband items require Streetwise to roll.
Additionally, when equipped to a living Rixixi, treats that Rixixi as a Ghost in all ways except for genetics.
Lucky Charm - Special Ability: Dowsing Rod: The Ixi holding this charm, while mining, will also roll one extra item - rolled from items such as Water, Dairy, Seasonal Drinks, and other beverages! Full list here!
Lucky Charm - Special Ability: Feint: The Ixi holding this charm may reroll a single attack per battle, and choose the attack with the better roll. (Reply to combat roll to use this ability)
Lucky Charm - Special Ability: Gift Shop: The Ixi holding this charm will always roll more than one item in explorations [unless affected by radiation sickness]!
Lucky Charm - Special Ability: Trophy: The Ixi holding this charm will always roll at least one Uncommon or Rare item when hunting. If they roll a single item, that item cannot be common.
Mini Rooter - Companion Ability: Mini Rooter: 80% chance of digging up an uncommon item in Explorations and Mining. Returns 1 Root Vegetables on failed roll.
Mismi - Companion Ability: Mismi: This companion guarantees its companion Rixixi will never roll fewer than 2 items in Mining and Exploring!
Mymber - Companion Ability: Mymber: If Chasmjumping with a Rixixi with this companion equipped, the player can request Teleportation, Danger, or Eyeseeking*, and the Mymber will attempt to lead the Rixixi to one of the Chasmjumping rolls in the selected category - 25% success rate!
*Teleporation includes Pathway, Parsec, and Liminal.
Danger includes Altercation, Facedown, AlternateFatality, and Prepare For War.
Eyeseeking includes Euphoria and Event Horizon.
A Mymber can also, if equipped to a radiation sick Rixixi, nullify the effects of one of the Radiation Sicknesses so long as the Mymber stays equipped! (Please specify which radiation sickness will be nullified when applying companion)
Mymber Tome - Special Ability: Expert at Humming: The Ixi with this ability can understand Mymbers. Any instance of Mymbers interacting with the Ixi will be translated.
MymbMumbles Lookalike - 'Jim'/MymbMumbles Lookalike - 'Jinn' - Companion Ability:
Counts as a Mymber in any scenarios that specifically require one, such as needing a tutor for the Mymber Tome. MymbMumbles Lookalikes are Mymbers with an extra skill.
MymbMumbles Lookalike: If pictured in an activity with another Rixixi with a MymbMumbles Lookalike, both Rixixi have their stats doubled. If pictured in a Training Mode image with another Rixixi with a MymbMumbles Lookalike, the EXP count is doubled for those Rixixi. This does work if there are more than two Rixixi in one image with MymbMumbles Lookalikes, but Rixixi in the image without MymbMumbles Lookalikes will not have the doubled stats/EXP effect!
Pack Cat - Companion Ability: Pack Cat: This companion can add on an entire additional roll (including seasonal item returns) to its companion Rixixi whenever the Rixixi it is attached to is depicted in an exploration or mining roll! The additional roll will be rolled with base stats rather than the Rixixi's stats, as the Pack Cat is picking up the items rather than the Rixixi.
Parasite Companion - Companion Ability - Parasite: When used in combat, the Parasite's Flytrap stops an opponent from moving for 2 turns. Any skills that require moving fail. Dodging is not possible.
Legendary Companion Second Ability:
Parasite: Biting You Biting You Biting You: Inflicts 2d10 Insect Damage on the opponent for 2-5 turns. Insect Damage will not add more turns while already active.
Rooter - Companion Ability: Rooter: 50% chance of digging up a rare item in Explorations and Mining. Returns 1 Root Vegetables on failed roll. Also adds a bonus +50 to Speed to its companion Rixixi in these activities. (OVERRIDES Radiation Sickness if present!)
Rootershell Armor Special Ability: Medium Armor: Automatically blocks damage in combat as long as equipped items are allowed - Damage blocked = [Defense/2]+[Attack/2]+1d100. IF your Attack is above 200, 10% chance to automatically inflict damage whenever attacked (Damage= d20+Attack/2).
Roothorne Squeaker - Companion Ability: Roothorne: As they are closely related to the Rooter, Roothornes have the original Rooter skill and an additional skill.
The Rixixi this Roothorne is equipped to gets double EXP when pictured with another ARPG species!
Roothornes count as Rooters for the Grand Old Rooter's ability.
Skydrake - Companion Ability: Skydrake: Adds a bonus +100 Speed in Exploring and Mining.
If the Rixixi has a Tunneldrake equipped, the Tunneldrake's return is doubled.
Legendary Companion Second Ability:
Skydrake: Death From Above: When Legendary Hunting, adds 15% Success chance to entire party. Retrieves one Rack of Ribs on a failed Legendary Hunt.
Snared Mymber- Companion Ability:
Counts as a Mymber in any scenarios that specifically require one, such as needing a tutor for the Mymber Tome. Snared Mymbers are Mymbers with an extra skill.
Snared Mymber: Can assist Rixixi companion in combat with the following skill:
Snowman - Companion Ability: Snowman: Blocks Temporary Conditions that can be considered 'exposure to the elements'. (Currently Cursed, Radiation Poisoning, and Vent Smog)
Legendary Companion Second Ability:
Snowman: Frostbite: Any Damage Over Time dealt by the companion Rixixi is doubled.
Spring's Messengerbird - Companion Ability: Spring's Messengerbird: Due to Advanced Bird Communication, if pictured with another Rixixi with a Spring's Messengerbird in any loot-dropping activity, the Rixixi it is attached to gets an additional roll (rolled at base stats rather than the Rixixi's stats)! The additional roll is only available to Rixixi in the activity piece with Spring's Messengerbird Companions.
Also acts as a version of the Floral Tea boon that can be activated on any roll the attached Rixixi is in: Use to reroll all items returned in a loot-dropping activity roll. (To use, head to claims. Add all items from the original roll into Add-Ons, link to the original roll, and state that you're using Spring's Messengerbird. The reroll will be previewed in a comment, you may then choose whichever roll you would like to keep.) Floral Tea cannot be used on this Rixixi as it would have the same effect.
Summertime Fogbee - Companion Ability: Summertime Fogbee: If Chasmjumping with a Rixixi with this companion equipped, the Rixixi will receive one jar of Honey on any one-day-cooldown results such as Nothing or YOU ARE NOT READY Event Horizons.
Also acts as a version of the Blue Raspberry Lemonade boon that can be activated on an Exploring, Hunting, or Mining roll the attached Rixixi is in: Your Fogbee will roll an extra item! (Item will be rolled from the pool of already-available items that your roll was drawn for - that is, the canon world for Explorations and the prey type for Hunting) Blue Raspberry Lemonade cannot be used on this Rixixi as it would have the same effect.
Tail Bags - Special Ability: Extra Storage: The Rixixi wearing this apparel came prepared and gets an entire additional roll in any loot-dropping activity! The additional roll will be rolled with base stats rather than the Rixixi's stats.
Taranan - Companion Ability: Taranan: Rolls 2 6-sided die. Doubles roll on doubles and sevens.
Legendary Companion Second Ability:
Taranan: Famous Last Words: When Legendary Hunting, rerolls a failed legendary hunt once.
Trinket Jar - Potpourri - Special Ability - Chef's Utility Belt: Can be used only once a week. When crafting in the Kitchen, link the Rixixi this skill is on to use this skill's weekly charge in place of all spices in a single Kitchen recipe. (Must be an existing recipe, not one being created.)
Trinket Jar - Sand Art - Special Ability - Jar of Art: This Rixixi and any others portrayed with them in Training Mode pieces get a +10 EXP!
Trophy - Companion Ability: Trophy: Adds 5 Seasonal Currency on top of the regular currency drop. This applies to every currency drop in the roll - main roll is affected, as are bonuses such as tail bags and pack cats.
Legendary Companion Second Ability:
Trophy: Covered By Roses: Can be used during combat to restore 25% of total HP. Has a 3-turn cooldown.
Tunneldrake - Companion Ability: Tunneldrake: This companion guarantees its companion Rixixi will never roll fewer than 2 items in Mining and Hunting (OVERRIDES Radiation Sickness if present!), and in those two activities will also bring back one bonus item!
Warren (content warnings: amalgamation, swarming, body horror) - Companion Ability: Warren: Tramples opponents in combat, ignoring armor.
Legendary Companion Second Ability:
Warren: There's A Fog Along The Horizon: 65% chance to dodge the enemy's next 3 moves, increased critical chance by 2 during duration of the move due to stealth. Cannot be used when already active. 40% chance to fail if used in succession. Must be activated to use, taking up a combat turn.
Warreneater - Companion Ability: Warreneater: Doubles hunting rolls.
Legendary Companion Second Ability:
Warreneater: Run Rabbit Run: If the Rixixi Companion fails a legendary hunt, the hunt will reroll as a Misc. Land Animal hunting at base stats.
Winterbringer Deer - Winterbringer Deer: The Rixixi with this companion cannot bring back less than 2 items in any loot-dropping activity if the piece has ice or snow portrayed.
Also automatically adds the Deepearth Brew boon to any roll: Doubles the quantity of all returned items in the activity, but will not double the return of items that were found due to the use of items that guarantee the return of a certain item, such as Glass Bottle and its ability to return 1x Water, 1x Science Kit, or 1x Deepearth Brew. Deepearth Brew cannot be used on this Rixixi as it would have the same effect.
Skills granted through other Story Modes or other methods.
Some combat-based skills, as shown in these lists, cause extra status effects. While the full effects of the skills that cause these may vary, some basic statuses exist in many skills, and as such, are described here: